Academia.eduAcademia.edu

AI-generated Abstract

This study investigates the bioarchaeological aspects of human remains found in mass graves at the Phaleron cemetery in Greece, which offers insights into societal perceptions of death and burial practices during the Archaic era. By analyzing various types of burials, including deviant and violent ones, the research aims to fill gaps in the understanding of how ancient Attic society dealt with death and its implications.

References (101)

  1. Malmström et al. 2009; Svensson et al. 2007; Yang et al. 1998. 105 Meyer & Kircher 2010.
  2. Günther et al. 2018; 2015.
  3. Meyer & Kircher 2010.
  4. Li & Durbin 2010. 109 Kircher 2012.
  5. Briggs et al. 2007; Hansen et al. 2001; Hofreiter et al. 2001; Orlando et al. 2011; Sawyer et al. 2012. 111 Skoglund et al. 2014. Bibliography
  6. Angel, J.L. 1975. 'Paleoecology, paleodemography and health' , in Population, ecology and social evolution, ed. S. Polgar, The Hague, 167-190. https://doi.org/10.1515/9783110815603.167
  7. Arbour, L. 2008. `Blunt force trauma´, in Skeletal trauma. Identification of injuries resulting from human rights abuse and armed conflict, ed. E.H. Kimmerle & J.P. Baraybar, Boca Raton, Florida, 151-195.
  8. Aufderheide, A.C. & C. Rodrígues-Martin 1998. The Cambridge encyclopedia of human paleopathology, Cambridge.
  9. Boehlau, J. 1887. 'Frühattische Vasen' , Jdl 2, 33-66.
  10. Borrini, M., P.P. Mariani, C. Murgia, C. Rodriguez & T.V. Tumbarello 2012. 'Contextual taphonomy. Superficial bone alterations as contextual indicators' , Journal of Biological Research 85:1, 217-219. https://doi.org/10.4081/jbr.2012.4115
  11. Briggs, A.W., U. Stenzel, P.L.F. Johnson, R.E. Green, J. Kelso, K. Prüfer, M. Meyer, J. Krause, M.T. Ronan, M. Lachmann & S. Pääbo 2007. 'Patterns of dam- age in genomic DNA sequences from a Neandertal' , Proceedings of the National Academy of Sciences of the United States of America 104, 14616-14621. https://doi.org/10.1073/pnas.0704665104
  12. Brothwell, D.R. 1972 2 . Digging up bones. The excavation, treat- ment and study of human skeletal remains, London.
  13. Buikstra, J.E. & D.H. Ubelaker, eds. 1994. Standards for data collection from human skeletal remains. Proceedings of a seminar at the Field Museum of Natural History organized by Jonathan Haas (Arkansas Archeological Survey Research Series, 44), Fayetteville, Arkansas.
  14. Burk, M.P. 2012. Forensic pathology of fractures and mecha- nisms of injury. Postmortem CT scanning, London. https://doi.org/10.1201/b11494
  15. Byers, S.N. 2008 3 . Introduction to forensic anthropology, Boston. https://doi.org/10.4324/9781315642031
  16. Calce, T. & T.L. Rogers 2007. 'Taphonomic changes of blunt force trauma. A preliminary study' , Journal of Forensic Science 52:3, 519-527. https://doi.org/10.1111/j.1556-4029.2007.00405.x Chryssoulaki, S. forthcoming. 'The excavations of Phaleron cemetery 2012-2017. An introduction' , in Rethink- ing Athens. Interdisciplinary approaches to the polis before the Persian Wars. Internationalen Workshop für junge Wissenschaftler vom 23.-24. Februar 2017 an der Ludwig-Maximilians-Universität München, eds.
  17. C. Graml, A. Doronzio & V. Capozzoli, Munich.
  18. Cole, G. & T. Waldron 2016. 'Purple staining of archaeo- logical human bone. An investigation of probable cause and implications for other tissues and artifacts' , Journal of Anthropology 2016, article ID 9479051. https://doi.org/10.1155/2016/9479051
  19. DiMaio V.J. & D. DiMaio 2001 2 . Forensic pathology, Boca Raton, Florida.
  20. Dimitriadou, E.M. 2012. Πρώιμη Αθήνα (1100-480 π.Χ.). Παρατηρήσεις στην οικιστική εξέλιξη και στα νεκροταφεία, Ph.D. thesis, National and Kapodistrian University of Athens.
  21. Dirkmaat, D.C., L.L. Cabo, S.D. Ousley & S.A. Symes 2008. 'New perspectives in forensic anthropology' , Yearbook of Physical Anthropology 51, 33-52. https://doi.org/10.1002/ajpa.20948
  22. Duday, H. 2009. The archaeology of the dead. Lectures in archaeothanatology (Studies in Funerary Archaeol- ogy, 3), Oxford.
  23. Dufton, A. & C. Fenwick 2012. 'Beyond the grave. Develop- ing new tools for medieval cemetery analysis at Vil- lamagna, Italy' , in Thinking beyond the tool. Archaeo- logical computing and the interpretive process (BAR-IS, 2344), eds. A. Chrysanthi, P. Murrieta Flores & C. Papadopoulos, Oxford, 155-167.
  24. Dupras, T.L. & J.J. Schultz 2013. 'Taphonomic bone staining and color changes in forensic contexts' , in Manual of forensic taphonomy, eds. J.T. Pokines & S.A. Symes, Boca Raton, Florida, 315-340. https://doi.org/10.1201/b15424-13
  25. Finegan, O. 2008. 'Case study. The interpretation of skeletal trauma resulting from injuries sustained prior to, and as a direct result of freefall' , in Skeletal trauma. Iden- tification of injuries resulting from human rights abuse and armed conflict, eds. E.H. Kimmerle & J.P. Baraybar, Boca Raton, Florida, 181-195.
  26. Galloway, A. & V.L. Wedel 2014 2 . 'Bones of the skull, the dentition and osseus structures of the throat' , in Broken bones. Anthropological analysis of blunt force trauma, eds. V.L. Wedel & A. Galloway, Springfield, Illinois, 133-160.
  27. Galloway, A., L. Zephro & V.L. Wedel 2014 2 . 'Diagnostic criteria for the determination and classification of timing and fracture mechanism' , in Broken bones. An- thropological analysis of blunt force trauma, eds. V.L. Wedel & A. Galloway, Springfield, Illinois, 47-58.
  28. Goodman, A.H. & J.C. Rose 1990. 'Assessment of systemic physiological perturbations from dental enamel hypo- plasias and associated histological structures' , Ameri- can Journal of Physical Anthropology 33, 59-110. https://doi.org/10.1002/ajpa.1330330506
  29. Goodman, A., G. Armelagos & J. Rose 1980. 'Enamel hypo- plasias as indicators of stress in three prehistoric popu- lations from Illinois' , Human Biology 52:3, 515-528.
  30. Günther, T., H. Malmström, E.M. Svensson, A. Omrak, F. Sánchez-Quinto, G.M. Kılınç, M. Krzewińska, G. Eriksson, M. Fraser, H. Edlund, A.R. Munters, A. Coutinho, L.G. Simões, M. Vicente, A. Sjölander, B. Jansen Sellevold, R. Jørgensen, P. Claes, M.D. Shriver, C. Valdiosera, M.G. Netea, J. Apel, K. Lidén, B. Skar, J. Storå, A. Götherström & M. Jakobsson 2018. 'Population genomics of Mesolith- ic Scandinavia. Investigating early postglacial migration routes and high-latitude adaptation' , PLOS Biology 16, article ID e2003703. https://doi.org/10.1371/journal.pbio.2003703
  31. Günther, T., C. Valdiosera, H. Malmström, I. Ureña, R. Rodriguez-Varela, Ó.O. Sverrisdóttir, E.A. Daska- laki, P. Skoglund, T. Naidoo, E.M. Svensson, J.M. Bermúdez de Castro, E. Carbonell, M. Dunn, J. Storå, E. Iriarte, J.L. Arsuaga, J.-M. Carretero, A. Götherström & M. Jakobsson 2015. 'Ancient genomes link early farmers from Atapuerca in Spain to modern-day Basques' , Proceedings of the National Academy of Sciences 112, 11917-11922. https://doi.org/10.1073/pnas.1509851112
  32. Guyomarc'h, P., M. Campagna-Vaillancourt, C. Kremer & A. Sauvageau 2010. 'Discrimination of falls and blows in blunt head trauma. A multi-criteria ap- proach' , Journal of Forensic Sciences 55:2, 423-427. https://doi.org/10.1111/j.1556-4029.2009.01310.x
  33. Hansen, A., E. Willerslev, C. Wiuf, T. Mourier, P. Arctander 2001. 'Statistical evidence for miscoding lesions in ancient DNA templates' , Molecular Biology and Evo- lution 18:2, 262-265. https://doi.org/10.1093/oxfordjournals.molbev.a003800
  34. Hillson, S. 1996. Dental anthropology, Cambridge. https://doi.org/10.1017/CBO9781139170697
  35. Hofreiter, M., V. Jaenicke, D. Serre, A. von Haeseler, S. Pääbo 2001. 'DNA sequences from multiple amplifications reveal artifacts induced by cytosine deamination in ancient DNA' , Nucleic Acids Research 29, 4793-4799. https://doi.org/10.1093/nar/29.23.4793
  36. Keramopoulos, A. 1923. Αποτυμπανισμός. Συμβολή αρχαιολογική εις την ιστορίαν του Ποινικού Δικαίου και την Λαογραφία, Athens.
  37. Kimmerle, E.H. & J.P. Baraybar, eds. 2008. Skeletal trauma. Identification of injuries resulting from human rights abuse and armed conflict, Boca Raton, Florida. https://doi.org/10.1201/9781420009118
  38. Kircher, M. 2012. ' Analysis of high-throughput ancient DNA sequencing data' , in Methods in molecular biology, eds. B. Shapiro & M. Hofreiter, New York, 197-228. https://doi.org/10.1007/978-1-61779-516-9_23
  39. Knüsel, C.J. 2014. 'Crouching in fear. Terms of engagement for funerary remains' , Journal of Social Archaeology 14, 26-58. https://doi.org/10.1177/1469605313518869
  40. Knüsel, C.J. & J. Robb 2016. 'Funerary taphonomy. An over- view of goals and methods' , JAS Reports 10, 655-673. https://doi.org/10.1016/j.jasrep.2016.05.031
  41. Kourouniotis, K. 1911. 'Εξ Αττικής. Α΄ Ανασκαφαί Παλαιού Φαλήρου' , AEphem 50, 246-251.
  42. Kranioti, E.F. 2015. 'Forensic investigation of cranial injuries due to blunt force trauma. Current best practice' , Research and Reports in Forensic Medical Science 2015:5, 27-35. https://doi.org/10.2147/rrfms.s70423
  43. Kremer, C., S. Racette, C.-A. Dionne & A. Sauvageau 2008. 'Discrimination of falls and blows in blunt force head trauma. Systematic study of the hat brim line rule in relation to skull fractures' , Journal of Forensic Sciences 53:3, 716-719. https://doi.org/10.1111/j.1556-4029.2008.00725.x
  44. Krogman, W.M. 1978. The human skeleton in forensic medi- cine, Springfield, Illinois.
  45. Krovitz, G.E. & P. Shipman 2007. 'Taphonomy of immature hominid skulls and the Taung, Mojokerto, and Herto specimens' , in Breathing life into fossils. Taphonomic studies in honor of C.K. (Bob) Brain (Stone Age Insti- tute Publication Series, 2), eds. T.R. Pickering, K. Schick & N. Toth, Gosport, Indiana, 207-232.
  46. Larsen, C.P. 1997 1 . Bioarchaeology. Interpreting behavior from the human skeleton (Cambridge Studies in Biological Anthropology, 21), Cambridge.
  47. Larsen, C.P. 2015 2 . Bioarchaeology. Interpreting behavior from the human skeleton (Cambridge Studies in Biological and Evolutionary Anthropology, 69), Cambridge.
  48. Li, H. & R. Durbin 2010. 'Fast and accurate long-read align- ment with Burrows-Wheeler transform' , Bioinfor- matics 26, 589-595. https://doi.org/10.1093/bioinformatics/btp698
  49. Lieverse, A.R. 1999. 'Diet and the aetiology of dental calculus' , International Journal of Osteoarchaeology 9:4, 219-232. https://doi.org/10.1002/(sici)1099- 1212(199907/08)9:4<219::aid-oa475>3.0.co;2-v
  50. Lillie, M.C. 1996. 'Mesolithic and Neolithic populations of Ukraine. Indications of diet from dental pathology' , CurrAnthr 37:1, 135-142. https://doi.org/10.1086/204479
  51. Liston, M.A. & L.P. Day 2009. 'It does take a brain surgeon. A successful trepanation from Kavousi, Crete' , in New directions in the skeletal biology of Greece (Occasional Wiener Laboratory Series, 1), eds. L.A. Schepartz, S.C. Fox & C. Bourbou, Princeton, New Jersey, 57-73.
  52. Lovell, N.C. 1997. 'Trauma analysis in paleopathology' , Year- book of Physical Anthropology 40, 139-170. https://doi.org/10.1002/(sici)1096- 8644(1997)25+<139::aid-ajpa6>3.0.co;2-#
  53. Lukacs, J.R. 2007. 'Dental trauma and antemortem tooth loss in prehistoric Canary Islanders. Prevalence and contributing factors' , International Journal of Osteoar- chaeology 17:2, 157-173. https://doi.org/10.1002/oa.864
  54. Lyman, L. 2010. 'What taphonomy is, what it isn't, and why taphonomists should care about the difference' , Jour- nal of Taphonomy 8:1, 1-16.
  55. Malmström, H., M.T. Gilbert, M.G. Thomas, M. Brand- ström, J. Storå, P. Molnar, P.K. Andersen, C. Ben- dixen, G. Holmlund, A. Götherström & E. Willerslev 2009. 'Ancient DNA reveals lack of continuity be- tween Neolithic hunter-gatherers and contemporary Scandinavians' , Current Biology 19, 1758-1762. https://doi.org/10.1016/j.cub.2009.09.017
  56. Mann, R.W. & D.R. Hunt 2005 2 . Photographic regional atlas of bone disease. A guide to pathologic and normal variation in the human skeleton, Springfield, Illinois.
  57. Maples, W.R. 1986. 'Trauma analysis by the forensic anthro- pologist' , in Forensic osteology. Advances in the identifi- cation of human remains, ed. K.J. Reichs, Springfield, Illinois, 218-228.
  58. Meyer, M. & M. Kircher 2010. 'Illumina sequencing library preparation for highly multiplexed target capture and sequencing' , Cold Spring Harbor Protocols 2010, article ID pdb.prot5448. https://doi.org/10.1101/pdb.prot5448
  59. Moritz, A.R. 1954 2 . The pathology of trauma, Philadelphia, Pennsylvania.
  60. Morris, I. 1989. 'Attitudes toward death in Archaic Greece' , ClAnt 8:2, 296-320. https://doi.org/10.2307/25010911
  61. Morris, I. 1992. Death-ritual and social structure in Classical Antiquity, Cambridge. https://doi.org/10.1017/CBO9780511611728
  62. Nawrocki, S.P., J.E. Pless, D.A. Hawley & S.A. Wagner 1997. 'Fluvial transport of human crania' , in Forensic tapho- nomy. The postmortem fate of human remains, eds.
  63. W.D. Haglund & M.H. Sorg, Boca Raton, Florida, 529-548. https://doi.org/10.1201/9781439821923.sec5
  64. Orlando, L., A. Ginolhac, M. Raghavan, J. Vilstrup, M. Ras- mussen, K. Magnussen, K.E. Steinmann, P. Kapranov, J.F. Thompson, G. Zazula, D. Froese, I. Moltke, B. Shapiro, M. Hofreiter, K.A. Al-Rasheid, M.T. Gilbert & E. Willerslev 2011. 'True single-molecule DNA sequencing of a Pleistocene horse bone' , Ge- nome Research 21, 1705-1719. https://doi.org/10.1101/gr.122747.111
  65. Ortner, D.J. 2003 2 . Identification of pathological conditions in human skeletal remains, Amsterdam. https://doi.org/10.1016/B978-0-12-528628-2.X5037-6
  66. Ortner, D.J. & G.J. Putschar 1985 1 . Identification of pathologi- cal conditions in human skeletal remains (Smithsonian Contributions to Anthropology, 28), Washington.
  67. Parker, R. 1983. Miasma. Pollution and purification in early Greek religion, Oxford.
  68. Pelekidis, S. 1916. 'Ανασκαφή Παλαιού Φαλήρου' , ADelt B2, 13-64.
  69. Philippson, A. & E. Kirsten 1952. Die griechischen Land- schaften 1:3. Attika und Megaris, Frankfurt.
  70. Pink, C.M. 2016. 'Forensic ancestry assessment using cranial nonmetric traits traditionally applied to biologi- cal distance studies' , in Biological distance analysis. Forensic and bioarchaeological perspectives, eds. M.A. Pilloud & J.T. Hefner, London, 213-230. https://doi.org/10.1016/b978-0-12-801966-5.00011-1
  71. Rodríguez-Martin, C. 2006. 'Identification and differential diagnosis of traumatic lesions of the skeleton' , in Forensic anthropology and medicine. Complementary sciences from recovery to cause of death, eds. A. Schmitt, E. Cunha & J. Pinheiro, Totowa, New Jersey, 197-221. https://doi.org/10.1007/978-1-59745-099-7_8
  72. Rogers, J. & T. Waldron 1995. A field guide to joint disease in archaeology, Chichester & New York.
  73. Rogers, J., L. Shepstone & P. Dieppe 1997. 'Bone formers. Osteophyte and enthesophyte formation are posi- tively associated' , Annals of the Rheumatic Diseases 56:2, 85-90. https://doi.org/10.1136/ard.56.2.85
  74. Sauer, N.J. 1998. 'The timing of injuries and manner of death. Distinguishing between antemortem, perimor- tem and postmortem trauma' , in Forensic osteology. Advances in the identification of human remains, ed. K.J. Reichs, Springfield, Illinois, 321-332.
  75. Sawyer, S., J. Krause, K. Guschanski, V. Savolainen & S. Pääbo 2012. 'Temporal patterns of nucleotide mis- incorporations and DNA fragmentation in ancient DNA' , PLoS ONE 7, article ID e34131. https://doi.org/10.1371/journal.pone.0034131
  76. Scheuer, L. & S.M. Black 2000. Developmental juvenile osteol- ogy, San Diego, California. https://doi.org/10.1016/b978-0-12-624000-9.x5000-x
  77. Schmitt, H., E. Dubljanin, S. Schneider & M. Schiltenwolf 2004. 'Radiographic changes in the lumbar spine in former élite athletes' , Spine 29:22, 2554-2559.
  78. Skoglund, P., B.H. Northoff, M.V. Shunkov, A.P. Dere- vianko, S. Pääbo, J. Krause & M. Jakobsson 2014. 'Separating endogenous ancient DNA from modern- day contamination in a Siberian Neandertal' , Proceed- ings of the National Academy of Sciences of the United States of America 111, 2229-2234. https://doi.org/10.1073/pnas.1318934111
  79. Skoglund, P., J. Storå, A. Götherström & M. Jakobsson 2013. ' Accurate sex identification of ancient human remains using DNA shotgun sequencing' , JAS 40, 4477-4482. https://dx.doi.org/10.1016/j.jas.2013.07.004
  80. Smith, C. 1884. 'Pyxis. Herakles and Geryon' , JHS 5, 176-184. https://doi.org/10.2307/623702
  81. Spitz, W.U. & D.J. Spitz, eds. 1993 3 . Spitz and Fisher's medico- legal investigation of death. Guidelines for the application of pathology to crime investigation, Springfield, Illinois.
  82. Spradley, M.K. & R.L. Jantz 2011. 'Sex estimations in foren- sic anthropology. Skull versus postcranial elements' , Journal of Forensic Sciences 56:2, 289-296. https://doi.org/10.1111/j.1556-4029.2010.01635.x
  83. Stewart, T.D. 1979. Essentials of forensic anthropology espe- cially as developed in the United States, Springfield, Illinois.
  84. Stuart-Macadam, P. 1992. 'Anemia in past human popula- tions' , in Diet, demography and disease. Changing perspectives on anemia, eds. P. Stuart-Macadam & S. Kent, New York, 151-170.
  85. Svensson, E.M., C. Anderung, J. Baubliene, P. Persson, H. Malmström, C. Smith, M. Vretemark, L. Daugno- ra & A. Götherström 2007. 'Tracing genetic change over time using nuclear SNPs in ancient and modern cattle' , Animal Genetics 38:4, 378-383. https://doi.org/10.1111/j.1365-2052.2007.01620.x
  86. Symes, S.A., E.N. L' Abbé, K.E. Stull, M. Lacroix & J.T. Pokines 2014. 'Taphonomy and the timing of bone fractures in trauma analysis' , in Manual of foren- sic taphonomy, eds. J.T. Pokines & S.A. Symes, Boca Raton, Florida, 341-366. https://doi.org/10.1201/b15424-14
  87. Ta' Ala, S.C., G.E. Berg. & K. Haden 2006. 'Blunt force trauma in the Cambodian Killing Fields' , Journal of Forensic Sciences 51:5, 996-1001. https://doi.org/10.1111/j.1556-4029.2006.00219.x Thapliyal, G.K., R. Sinha, P.S. Menon & A. Chakranarayan 2008. 'Management of mandibular fractures' , Medical Journal Armed Forces India 64:3, 218-220. https://doi.org/10.1016/s0377-1237(08)80096-2
  88. Trotter, M. 1970. 'Estimation in stature from intact long limb bones' , in Personal identification in mass disasters. Report of a seminar held in Washington, D.C., 9-11 December 1968, by arrangement between the Sup- port Services of the Department of the Army and the Smithsonian Institution, ed. T.D. Stewart, Washing- ton, 71-84.
  89. Ubelaker, D.H. 1989 2 . Human skeletal remains. Excavation, analysis, interpretation (Manuals on Archaeology, 2), Washington, D.C.
  90. Ubelaker, D.H. & K.M. Montaperto 2014. 'Trauma inter- pretation in the context of biological anthropology' , in The Routledge handbook of the bioarchaeology of human conflict, eds. C. Knüsel & M.J. Smith, Oxford & New York, 25-38. https://doi.org/10.4324/9781315883366.ch2
  91. Unnewher, M., C. Homann, P.F. Schmidt, P. Sotony, G. Fischer, B. Brinkmann, T. Bajanowski & A. DuChesne 2003. 'Fracture properties of the human mandible' , International Journal of Legal Medicine 117:6, 326-330.
  92. Valavanis, P. 2010. 'Αθήνα' , in Αρχαία Αθήνα και Αττική. Ιστορική Τοπογραφία του Άστεως και της Χώρας, ed. A.I. Vlachopoulos, Athens, 40-61.
  93. Vianello, D., F. Sevini, G. Castellani, L. Lomartire, M. Capri & C. Franceschi 2013. 'HAPLOFIND. A new method for high-throughput mtDNA haplogroup assignment' , Human Mutation 34, 1189-1194. https://doi.org/10.1002/humu.22356
  94. Walker, P.L. 1995. 'Problems of preservation and sexism in sexing. Some lessons from historical collections for palaeodemographers' , in Grave reflections. Portraying the past through cemetery studies, eds. S.R. Saunders & A. Herring, Toronto, 31-47.
  95. Walker, P.L., R.R. Bathurst, R. Richman, T. Gjerdrum & V.A. Andrushko 2009. 'The causes of porotic hyperostosis and cribra orbitalia. A reappraisal of the iron-deficiency-anemia hypothesis' , American Journal of Physical Anthropology 139:2, 109-125. https://doi.org/10.1002/ajpa.21031
  96. Weiss, E. 2006. 'Osteoarthritis and body mass' , JAS 33:5, 690-695. https://doi.org/10.1016/j.jas.2005.10.003
  97. Weiss, E. & R. Jurmain 2007. 'Osteoarthritis revisited. A contemporary review of aetiology' , International Journal of Osteoarchaeology 17:5, 437-450. https://doi.org/10.1002/oa.889
  98. Weston, D.A. 2012. 'Nonspecific infections in palaeopathol- ogy. Interpreting periosteal reactions' , in A companion to paleopathology (Blackwell Companions to Anthro- pology, 14), ed. A.L. Grauer, Chichester, West Sussex & Malden, Massachussetts, 492-512. https://doi.org/10.1002/9781444345940.ch27
  99. White, T.D. 1991. Human osteology, San Diego.
  100. White, T.D., M.T. Black & P.A. Folkens 2012 3 . Human osteology, Amsterdam & London. https://doi.org/10.1016/C2009-0-03221-8
  101. Yang, D.Y., B. Eng, J.S. Waye, J.C. Dudar & S.R. Saunders 1998. 'Technical note. Improved DNA extraction from ancient bones using silica-based spin columns' , American Journal of Physical Anthropology 105:4, 539-543. https://doi.org/10.1002/(sici)1096- 8644(199804)105:4<539::aid-ajpa10>3.3.co;2-2